We were asking
How much can an early, inventory-preserving rearrangement alter the future before it alters the organism-level developmental programme?
An exact local rearrangement after q8 opened increasingly different microscopic trajectories without broadly moving formation timing or adult phenotype, which remained strongly tied to organism identity.
How much can an early, inventory-preserving rearrangement alter the future before it alters the organism-level developmental programme?
The microscopic paths separated strongly, while timing and coarse adult phenotype remained comparatively canalized; robustness here coexists with substantial causal freedom.
Which pre-form measurement predicts whether that freedom will later contract around transition timing?